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Committee Composition

Compensation Committee Audit Committee Nominating and Governance Committee
Independent Directors
Daniel Becker, M.D., Ph.D. Compensation Committee Audit Committee
Michael Gladstone Audit Committee Nominating and Governance Committee
Natalie Holles Compensation Committee Nominating and Governance Committee
John Josey, Ph.D., M.B.A. Compensation Committee Nominating and Governance Committee
Saira Ramasastry, M.S., M.Phil. Audit Committee
Non-Independent Directors
Jeremy Bender, Ph.D., M.B.A.
Julie Grant, M. Phil., M.B.A.
= Chair = Member = Chair of the Board = Lead Director